" commented on your image on the <a href='%{data_object_link}'>Encyclopedia of Life site</a>: %{comment_body}"
Looks like this item has a leading space. These should be avoided. Please check for other occurrences.
This is a patchwork message and needs to be fixed... I'm worried about keeping the same key when adding a variable, though, so I'm going to rename it as well.
Thanks.